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EDITOR
AARON J. LADMAN
DEPARTMENT OF ANATOMY, MAIL STOP 506
HAHNEMANN MEDICAL COLLEGE and HOSPITAL
230 NORTH BROAD STREET
PHILADELPHIA, PENNSYLVANIA 19102
ASSOCIATE EDITORS
G. ADOLPH ACKERMAN
Ohio State University
EDWIN S. BOATMAN
University of Washington
DALE E. BOCKMAN
Medical College of Georgia
ROBERT P. BOLENDER
University of Washington
ROBERT S. BRESSLER
Mount Sinai School of Medicine
RUTH ELLEN BULGER
The University of Texas
at Houston
NIKOLAJS CAUNA
University of Pittsburgh
RICHARD E. COGGESHALL
The University of Texas
Medical Branch a t Galveston
WILLIAM G. DAIL, JR.
The University of New Mexico
MARTIN DYM
Georgetown University
RICHARD H. EDMONDS
Albany Medical College
RICHARD A. ELLIS
Brown University
JOHN F. FALLON
University of Wisconsin
Madison
TIMOTHY P. FITZHARRIS
The Medical University of
South Carolina
CHARLES J. FLICKINGER
University of Virginia
DANIEL S. FRIEND
University of California
San Francisco
JAMES L. MATTHEWS
Baylor College of Dentistry
Dallas
BRYCE L. MUNGER
Pennsylvania State University
Hershey
WILLIAM B. NEAVES
The University of Texas
at Dallas
MARIAN R. NEUTRA
Harvard Medical School
CHARLES C.C. OMORCHOE
Loyola University
HELEN A. PADYKULA
The University of
Massachusetts
DAVID P. PENNEY
University of Rochester
E L I 0 RAVIOLA
Harvard Medical School
THOMAS S. REESE
National Institutes of
Health
PAUL J. REIER
University of Maryland
JACK ROSENBLUTH
New York University
WEBSTER S.S. J E E
LINDA SALAND
University of Utah
The University of New Mexico
ERNST KALLENBACH
GARY SCHOENWOLF
University of Florida
University of Utah
Gainesville
ARLEN R. SEVERSON
GORDON I. KAYE
University of Minnesota
Albany Medical College
J O ANNE V. SIMSON
ROBERT 0. KELLEY
Medical Universitv of
The University of New Mexico
South caro1ina
BARRY F. KING
MORRIS SMITHBERG
University of California
University of Minnesota
Davis
Medical School
RAYMOND F. GASSER
Louisiana State University
New Orleans
MICHAEL D. GERSHON
Columbia University
LAUREL E. GLASS
University of California
San Francisco
WILLIAM J. GONYEA
The University of Texas
at Dallas
DAMON C. HERBERT
The University of Texas
at San Antonio
ROBERT HIKIDA
Ohio University
JAMES W. HINDS,
Boston University
Medical Center
DAVID E. HINTON
West Virginia University
CLAIRE HUCKINS
Baylor College of Medicine
Houston
SUSUMU I T 0
Harvard Medical School
~~~
MIKEL H. SNOW
University of Southern
California
SERGE1 P. SOROKIN
Harvard School of
Public Health
A.K. SZAKAL
Medical College of Virginia
BERNARD TANDLER
Case Western
Reserve University
VIRGINIA M. TENNYSON
Columbia University
ROBERT J. TOMANEK
University of Iowa
JERRY S. TRIER
Peter Bent Brigham Hospital
Boston
JOHN A. TROTTER
The University of New Mexico
HERSHEY WARSHAWSKY
McGill University
ROBERT E. WATERMAN
The University of
New Mexico
FREDERICK WEZEMAN
Loyola University
School of Medicine
HARRY H. WILCOX
University of Tennessee
Medical Units
MARY C. WILLIAMS
University of California
San Francisco
RICHARD L. WOOD
University of Southern
California
_______.______
Manuscripts and illustrations intended for publication in The Anatomical Record should be sent to Dr. Aaron J. Ladman,
Department of Anatomy, Mail Stop 506, Hahnemann Medical College and Hospital, 230 North Broad Street, Philadelphia,
Pennsylvania 19102. Please read notice to contributors on inside back cover page for preparation instructions.
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Officersof the American Association of Anatomists for 1981-82: Elizabeth D. Hay, president; John E. Pauly, president-elect: Sanford L. Palay,
president-emeritus; Aaron J. Ladman, first vice-president: Richard P. Bunge, secondvice-president; M. Roy Schwarz,programsecretary;William
P. Jollie, secretary-treasurer.
.
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Thecodeat thebottomofthefirstpageofeacharticleinthisjournalindicates thecopyright owner'sconsent thatcopiesofthearticlemaybemade
within thelimitsof Sections 107 or 108of theU.S. Copyright Law. Forcopyingheyond theselimits, except asnoted below, consentisgivenon the
condition that the copier pay the stated per-copy fee through the Copyright Clearance Center, Inc., 21 Congress Street, Salem, MA 01970. This
consent does not extend to other kinds of copying not permitted by law, such as copying for general distribution, advertising or promotional
purposes, creating new collective works, or for resale.
THE ANATOMICAL RECORD (ISSN0003-276X)official organofthe American Associationof Anatomists, isowned hyThe Wistar Instituteof
AnatomyandBiologyandpublishedmonthlyby AlanR. Liss,Inc., 150FifthAvenue,NewYork,NY 10011.For 1982,subscriptionpriceis$240in
US.;$273inEurope, theMiddleEast, and Africa; and$264inallothercountries. (SubscriptionsgoingtoEurope, theMiddleEast, and Africawill
be sent air freight to Europe and mailed from there.) Payment must be made in U.S. dollars or at the current rate of exchange. For payment not
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Duplicate copies cannot be sent to replace issues not delivered because of failure to notify publisher of change of address. Exclusive agents in
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change of address to Alan R. Liss, Inc., 150 Fifth Avenue, New York, N.Y. 10011. Indexed by: Biological Abstracts, Chemical Abstracts, Current
ContentsiLife Sciences-Social Science Index, Excerpta Medica, Index Medicus, and Institute of Scientific Information-Soviet Union.
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Send Subscriptions and Business Correspondence to
ALAN R. LISS, INC. 150 FIFTH AVENUE, NEW YORK, NY 10011 (212) 741-2515
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