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Aaron J. Ladman, Editor
Department of Anatomy Mail Stop 408
Hahnemann University Broad & Vine Philadelphia, Pennsylvania 19102
G. Adolph Ackerman
Ohio State liniversity
Peter S. Amenta
Robert W"od Johnson
Medical School
J o h n V. Basmajian
Chrdokc Rehabilitation Center
>McMastcrI Jniversi t y
D a l e E. B o c k m a n
Medical College of Georgia
Robert P. B o l e n d e r
h i v c r s i t y o f Wdshington
Robert S. Bressler
New York <:allege of Podiatric
Ruth EUen Bulger
Institute of Medicine
Washington, D <:.
Margaret R. Byers
Cniversity of Washington
Edward C. Carlson
lhiversity of North Dakota
Nikolajs Cauna
llniversity of Pittsburgh
D o n a l d J. Donaldson
Webster S.S. J e e
University of Tennessee
University (if CJtah
Harvard Medical School
Ernst Kallenbach
Thomas S. R e e s e
Martin Dym
Georgetown University
University of Florida
Gordon 1. Kaye
Albany Medical College
Robert 0. KeUey
T h e University of New Mexico
Barry F. King
University of California
National Institutes of Health
Paul J. Reier
IJniversity of Florida
Stanley L. Erlandsen
University of Minnesota
Medical School
Richard A. Ellis
Brown 1Jniversity
J o h n F. Fallon
University of Wisconsin
D o n a l d A. Fischman
Cornell Liniversity Medical
Dr. Jef€rey T. Laitman
Charles J. Flickinger
llniversity of Virginia
Daniel S. Friend
llniversity of California
San Francisco
Geraldine F. Gauthier
T h e University of Massachusetts
Michael D . G e r s h o n
Columbia University
B e n R. Clower
Jack L. Haar
llnivcrsity of Mississippi
Medical College of Virginia
William G. Dail, Jr.
T h e l'nivcrsity o f New Mexico
D e n n i s M. D e Pace
Hahncmann 1:niversity
Richard D . Dey
West Virginia 1:niversity
New York llniversity
Liida Saland
Health Science Center
at Dallas
I'niversity o f Galifornia
San Francisco
Jack Rosenbluth
T h e University of New Mexico
Gary S c h o e n w o l f
University of Utah
Arlen R. Severson
University of Minnesota
!A)yola llniversity
School of Medicine
Gerald R. C u n h a
Lynn J. RomreU
llniversity o f Floridl
Medical University of
South Carolina
J o h n Clancy, Jr.
T h e llniversity o f Texas
Mcdical Branch at Galveston
Gainesvi 1It.
Eric R. Lacy
William J . Gonyea
T h e University of Texas
Richard E. CoggeshaU
Elio Raviola
Duane E. Haines
'University of Mississippi
D a m o n C. Herbert
T h e Miversity of Texas
at San Antonio
J o h n C. Herr
University of Virginia
Robert Hiklda
Ohio University
David E. H i n t o n
University of California
Manuscripts and illustrations intended for publication in The
Anatomical Record should be sent to Dr. Aaron J. Ladman,
Department of Andtomy, Mail Stop 408, Hahnemann University,
Broad & Vine, Philadelphia, Pennsylvania 19102. Please read
notice to contributors on inside back cover page for preparation
Officers o f the American Association of Anatomists for 1989-90:
Jerome Sutin, president; Roger R. Markwald, president emeritus;
Karen Hitchcock, president-elect; Jennifer H. LaVail, first vicepresident; Donald A. Fischman, second vice-president; Charles E.
Slonecker, program secretary; Marilyn Zimny, program secretaryelect; Robert D. Yates, secretary-treasurer.
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The paper o n which this journal is printed adheres to the
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Mount Sinai School of Medicine
New York
Thomas A. Marino
Temple University
School of Medicine
Roger R. Markwald
Medical College of Wisconsin
George F. Martin
Ohio State University
James L. Matthews
Baylor College of Dentistry
Dr. S a m G. McClugage
Louisiana State University
New Orleans
Peter J. Sheridan
T h e University of Texas at
San Antonio
Mike1 H. Snow
University of Southern
Sergei P. Sorokin
Boston University
A.K. Szakal
Medical College of Virginia
Bernard Tandler
Harvard Medical School
Case Western
Reserve Lhiversity
Robert J, T o m a n e k
University of Iowa
Jerry S. Trier
Peter Bent Brigham Hospital
John A. Trotter
T h e 1Jniversity of New Mexico
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College of Medicine
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R. Peter Meyer
Hahnemann University
B r y c e L. Munger
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H e l e n A. Padykula
T h e University of
McGill University
T h e University of New Mexico
Frederick Wezeman
Inyola University of Chicago
David P. P e n n e y
Lawrence C. Zoller
llniversitv of Rochester
Boston llniversity
THE ANATOMICAL RECORD (ISSN 0003-276)3), an official
publication of the American Association o f Anatomists, is
owned by the Wistar Institute of Anatomy and Biology and
published monthly by Alan R. Liss, Inc., 41 East 11th Street,
New York, NY 10003. For 1989, Volumes 223-225, twelve
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